Boll/bəʊl/ Definition 1 of 2Pro Tip 1/3Plant Case Boll refers to the protective spherical case around seeds of certain plants. Look at how the bolls protect the seeds inside. Pro Tip 2/3Agricultural Term Most commonly used in agriculture; especially when discussing cotton or flax. During harvest, farmers check if the bolls have opened. Pro Tip 3/3Mature Boll A 'mature boll' indicates that the fruit case has ripened and seeds are ready. The field is full of mature cotton bolls.